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the formation of vegetable mould-第14章

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triangle was doubled up; with the apex and base left sticking out 
of the tube。  As the sides of the triangles were three inches in 
length; the result of their being drawn into a tube or into a 
burrow in different ways; may be conveniently divided into three 
groups:  those drawn in by the apex or within an inch of it; those 
drawn in by the base or within an inch of it; and those drawn in by 
any point in the middle inch。

In order to see how the triangles would be seized by worms; some in 
a damp state were given to worms kept in confinement。  They were 
seized in three different manners in the case of both the narrow 
and broad triangles:  viz。; by the margin; by one of the three 
angles; which was often completely engulfed in their mouths; and 
lastly; by suction applied to any part of the flat surface。  If 
lines parallel to the base and an inch apart; are drawn across a 
triangle with the sides three inches in length; it will be divided 
into three parts of equal length。  Now if worms seized 
indifferently by chance any part; they would assuredly seize on the 
basal part or division far oftener than on either of the two other 
divisions。  For the area of the basal to the apical part is as 5 to 
1; so that the chance of the former being drawn into a burrow by 
suction; will be as 5 to 1; compared with the apical part。  The 
base offers two angles and the apex only one; so that the former 
would have twice as good a chance (independently of the size of the 
angles) of being engulfed in a worm's mouth; as would the apex。  It 
should; however; be stated that the apical angle is not often 
seized by worms; the margin at a little distance on either side 
being preferred。  I judge of this from having found in 40 out of 46 
cases in which triangles had been drawn into burrows by their 
apical ends; that the tip had been doubled back within the burrow 
for a length of between 1/20 of an inch and 1 inch。  Lastly; the 
proportion between the margins of the basal and apical parts is as 
3 to 2 for the broad; and 2。5 to 2 for the narrow triangles。  From 
these several considerations it might certainly have been expected; 
supposing that worms seized hold of the triangles by chance; that a 
considerably larger proportion would have been dragged into the 
burrows by the basal than by the apical part; but we shall 
immediately see how different was the result。

Triangles of the above specified sizes were scattered on the ground 
in many places and on many successive nights near worm…burrows; 
from which the leaves; petioles; twigs; &c。; with which they had 
been plugged; were removed。  Altogether 303 triangles were drawn by 
worms into their burrows:  12 others were drawn in by both ends; 
but as it was impossible to judge by which end they had been first 
seized; these are excluded。  Of the 303; 62 per cent。 had been 
drawn in by the apex (using this term for all drawn in by the 
apical part; one inch in length); 15 per cent。 by the middle; and 
23 per cent。 by the basal part。  If they had been drawn 
indifferently by any point; the proportion for the apical; middle 
and basal parts would have been 33。3 per cent。 for each; but; as we 
have just seen; it might have been expected that a much larger 
proportion would have been drawn in by the basal than by any other 
part。  As the case stands; nearly three times as many were drawn in 
by the apex as by the base。  If we consider the broad triangles by 
themselves; 59 per cent。 were drawn in by the apex; 25 per cent。 by 
the middle; and 16 per cent。 by the base。  Of the narrow triangles; 
65 per cent。 were drawn in by the apex; 14 per cent; by the middle; 
and 21 per cent。 by the base; so that here those drawn in by the 
apex were more than 3 times as many as those drawn in by the base。  
We may therefore conclude that the manner in which the triangles 
are drawn into the burrows is not a matter of chance。

In eight cases; two triangles had been drawn into the same burrow; 
and in seven of these cases; one had been drawn in by the apex and 
the other by the base。  This again indicates that the result is not 
determined by chance。  Worms appear sometimes to revolve in the act 
of drawing in the triangles; for five out of the whole lot had been 
wound into an irregular spire round the inside of the burrow。  
Worms kept in a warm room drew 63 triangles into their burrows; 
but; as in the case of the pine…leaves; they worked in a rather 
careless manner; for only 44 per cent。 were drawn in by the apex; 
22 per cent。 by the middle; and 33 per cent。 by the base。  In five 
cases; two triangles were drawn into the same burrow。

It may be suggested with much apparent probability that so large a 
proportion of the triangles were drawn in by the apex; not from the 
worms having selected this end as the most convenient for the 
purpose; but from having first tried in other ways and failed。  
This notion was countenanced by the manner in which worms in 
confinement were seen to drag about and drop the triangles; but 
then they were working carelessly。  I did not at first perceive the 
importance of this subject; but merely noticed that the bases of 
those triangles which had been drawn in by the apex; were generally 
clean and not crumpled。  The subject was afterwards attended to 
carefully。  In the first place several triangles which had been 
drawn in by the basal angles; or by the base; or a little above the 
base; and which were thus much crumpled and dirtied; were left for 
some hours in water and were then well shaken while immersed; but 
neither the dirt nor the creases were thus removed。  Only slight 
creases could be obliterated; even by pulling the wet triangles 
several times through my fingers。  Owing to the slime from the 
worms' bodies; the dirt was not easily washed off。  We may 
therefore conclude that if a triangle; before being dragged in by 
the apex; had been dragged into a burrow by its base with even a 
slight degree of force; the basal part would long retain its 
creases and remain dirty。  The condition of 89 triangles (65 narrow 
and 24 broad ones); which had been drawn in by the apex; was 
observed; and the bases of only 7 of them were at all creased; 
being at the same time generally dirty。  Of the 82 uncreased 
triangles; 14 were dirty at the base; but it does not follow from 
this fact that these had first been dragged towards the burrows by 
their bases; for the worms sometimes covered large portions of the 
triangles with slime; and these when dragged by the apex over the 
ground would be dirtied; and during rainy weather; the triangles 
were often dirtied over one whole side or over both sides。  If the 
worms had dragged the triangles to the mouths of their burrows by 
their bases; as often as by their apices; and had then perceived; 
without actually trying to draw them into the burrow; that the 
broader end was not well adapted for this purposeeven in this 
case a large proportion would probably have had their basal ends 
dirtied。  We may therefore inferimprobable as is the inference
that worms are able by some means to judge which is the best end by 
which to d
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